Journal Articles

• Kurtul, I., Eryaşar, A.R., Mutlu, T., Parker, B., Kaya, C., Bayçelebi, E., Haubrock, P.J., Tarkan, A.S., Britton, J.R., Sarı, H.M., Gedik, K., 2026. Microplastic ingestion in invasive mosquitofish (Gambusia holbrooki): a nationwide survey from Türkiye. Environmental Sciences Europe, 38 (1).
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• Minaz, M., Félix, L.M., Er, A., Köse, Ö., Ak, K., Alparslan, C., Serdar, O., Aydın, B., Kurtul, I., Hoseini, S.M., 2026. Evaluation of rainbow trout response to the optimal anesthetic concentration of citronella oil (Cymbopogon nardus) predicted by artificial neural network. Aquaculture, 613.
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• Haubrock, P.J., Soto, I., Cuthbert, R.N., Kurtul, I., Briski, E., 2025. Analysing factors underlying the reporting of established non-native species. Scientific Reports, 15 (1).
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• Kurtul, I., Haubrock, P.J., Kaya, C., Aksu, S., İlhan, A., Sari, H.M., Aygen, C., Soto, I., Parker, B., Britton, R., Tarkan, A.S., 2025. Spatial and seasonal patterns in fish assemblages of the Bakırçay river are associated with physicochemical and habitat parameters. Scientific Reports, 15 (1).
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• Kaya, C., Minaz, M., Şentürk Koca, Y., Oral Kaba, M., Kurtul, I., Aytan, Ü., 2025. Monitoring microplastics in a region with sensitive fish biodiversity: Tigris, Euphrates and Van Lake drainages in Irano-Anatolian hotspot. Environmental Sciences Europe, 37 (1).
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• Yoğurtçuoğlu, B., Tarkan, A.S., Ekmekçi, F.G., Kırankaya, Ş.G., Kurtul, I., Özcan, D., Karadal, O., Öndes, F., Türkmen, G., Sarı, H.M., Haubrock, P.J., Vilizzi, L., 2025. Turkish freshwaters as a case of rising invasion risk of aquarium fishes under climate change. Scientific Reports, 15 (1).
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• Haubrock, P.J., Novello, M., Abreo, N.A., Błońska, D., Sampaio Franco, A.C., Soto, I., Castaldelli, G., Katsanevakis, S., Kouba, A., Balzani, P., Kurtul, I., Tarkan, A.S., Britton, J.R., Briski, E., 2025. GFID: A Global Fish Invasion Database. Ecological Research, 40 (6).
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• Kurtul, I., Parker, B., Bayçelebi, E., Oral Kaba, M., Kaya, C., Mutlu, T., Akalın, S., Gedik, K., Haubrock, P.J., Tarkan, A.S., Britton, J.R., 2025. Historically low microplastic loads within white bream (Blicca bjoerkna) in the Marmara basin, Türkiye. Journal of Fish Biology, 107 (4), 1454-1459.
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• Haubrock, P.J., Novello, M., Abreo, N.A., Błońska, D., Sampaio Franco, A.C., Soto, I., Castaldelli, G., Katsanevakis, S., Kouba, A., Balzani, P., Kurtul, I., Tarkan, A.S., Briski, E., Britton, R., 2025. A Global Account of Established Non-Native Fish Species. Global Change Biology, 31 (8).
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• Kaba, M.O., Minaz, M., Kaya, C., Jouy, T., Kurtul, I., Aytan, Ü., 2025. Silent invaders of freshwater ecosystems: Unveiling the microplastic crisis threatening the world's largest soda lake (Lake Van, Türkiye). Journal of Great Lakes Research, 51 (4).
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• Ghazal, A., Paul, R., Tarkan, A.S., Kurtul, I., Pegg, J., Andreou, D., Britton, J.R., 2025. Domestication as the driver of lower chronic stress levels in fish in catch-and-release recreational fisheries and aquaculture versus wild conspecifics. Plos One, 20 (6 June).
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• Oficialdegui, F.J., Soto, I., Balzani, P., Cuthbert, R.N., Haubrock, P.J., Kourantidou, M., Manfrini, E., Tarkan, A.S., Kurtul, I., Macêdo, R.L., Musseau, C.L., Roy, K., Kouba, A., 2025. Non-Native Species in Aquaculture: Burgeoning Production and Environmental Sustainability Risks. Reviews in Aquaculture, 17 (3).
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• Haubrock, P.J., Parker, B., Błońska, D., Briski, E., Everts, T., Fernandez, R.D., Kouba, A., Kourantidou, M., Kurtul, I., Mammola, S., Musolin, D.L., Nuñez, M.A., Olden, J.D., Rasmussen, J.J., Renault, D., Russell, J.C., Sousa, R., Tarkan, A.S., Britton, J.R., 2025. Conceptual and ethical considerations in invasion science. Bioscience, 75 (4), 317-330.
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• Baycelebi, E., Kurtul, I., Onay, H., Kaya, C., Haubrock, P.J., Turan, D., 2025. Describing biological parameters of several endemic Salmo species from the Caspian Sea and Persian Gulf basins. Oceanological and Hydrobiological Studies, 54 (1), 21-32.
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• Kaya, C., Aksu, İ., Aksu, S., Kocabaş, M., Kurtul, I., Turan, D., 2025. Tracing Historical River Connectivity Through Freshwater Fish Assemblages: The Case of Göksu River and Lake Beyşehir. Ecohydrology, 18 (2).
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• Aksu, S., Emiroğlu, Ö., Balzani, P., Britton, J.R., Köse, E., Kurtul, I., Başkurt, S., Mol, O., Çınar, E., Haubrock, P.J., Oztopcu-Vatan, P., Tarkan, A.S., 2025. High trophic similarity between non-native common carp and gibel carp in Turkish freshwaters: Implications for management. Aquaculture and Fisheries, 10 (1), 141-148.
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• Kalaycı, G., Kurtul, I., Bayçelebi, E., Kaya, C., Turan, D., 2025. Integrated molecular and morphological evidence reveals a new Rhodeus species (Cyprinidae: Acheilognathidae) from the Turkish Aegean basin, with a redescription of Rhodeus meridionalis. Journal of Fish Biology.
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• Kaya, C., Saç, G., Hubert, N., Minaz, M., Kurtul, I., 2025. Post-2000 milestones in freshwater fish taxonomy: a comprehensive review of newly described species in Türkiye. Zoosystematics and Evolution, 101 (3), 1309-1324.
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• Haubrock, P.J., Everts, T., Abreo, N.A.S., Bojko, J., Deklerck, V., Dickey, J.W.E., Franco, A.C.S., García-Berthou, E., Katsanevakis, S., Kirichenko, N.I., Mammola, S., Nuñez, M.A., Parker, B., Scalera, R., Soto, I., Strubbe, D., Tarkan, A.S., Vilizzi, L., Adriaens, T., Balzani, P., Błońska, D., Briski, E., Brys, R., Burgess, A.L., Byers, J.E., Cano-Barbacil, C., Castaldelli, G., Dick, J.T.A., Dominguez Almela, V., Dimarco, R.D., Florencio, M., Kouba, A., Kourantidou, M., Kurtul, I., Martín-Forés, I., Morissette, O., Olden, J.D., Soares, B.E., Truszkowski, J., Verreycken, H., Kenis, M., Sousa, R., Britton, J.R., 2025. The impacts of biological invasions. Biological Reviews.
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• Tarkan, A.S., Emiroğlu, Ö., Aksu, S., Kurtul, I., Błońska, D., Bayçelebi, E., Soto, I., Chan, S.S., Haubrock, P.J., Bradshaw, C.J.A., 2024. Testing the Dispersal-Origin-Status-Impact (DOSI) scheme to prioritise non-native and translocated species management. Scientific Reports, 14 (1).
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• Haubrock, P.J., Soto, I., Kurtul, I., Kouba, A., 2024. Are long-term biomonitoring efforts overlooking crayfish in European rivers? Environmental Sciences Europe, 36 (1).
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• Kurtul, I., Haubrock, P.J., 2024. The need of centralized coordination to counter biological invasions in the European Union. Environmental Sciences Europe, 36 (1).
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• Haubrock, P.J., Kurtul, I., Kouba, A., 2024. Tracking aquatic non-native macroinvertebrate species in Germany using long-term data. Environmental Sciences Europe, 36 (1).
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• Onay, H., Kurtul, I., Kaya, C., Turan, D., Bayçelebi, E., 2024. Length-weight relations and condition factors of several endemic and native fish species from Anatolian freshwater. Oceanological and Hydrobiological Studies, 53 (4), 386-395.
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• Haubrock, P.J., Soto, I., Tarkan, A.S., Macêdo, R.L., Kouba, A., Cuthbert, R.N., Briski, E., Everts, T., Kurtul, I., 2024. Socioeconomic prerequisites determine national long-term biomonitoring efforts. Journal of Environmental Management, 370.
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• Haubrock, P.J., Kurtul, I., Macêdo, R.L., Mammola, S., Franco, A.C.S., Soto, I., 2024. Competency in invasion science: addressing stagnation challenges by promoting innovation and creative thinking. Environmental Management, 74 (5), 916-927.
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• Kurtul, I., 2024. The critical need for long-term biomonitoring: The case study of a major river system in Anatolia. Environmental Monitoring and Assessment, 196 (11).
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• Kurtul, I., Kaya, C., Onay, H., Bayçelebi, E., Turan, D., 2024. Length-weight relationships and condition factors of several endemic and native species of the genera Alburnoides, Alburnus and Squalius. Oceanological and Hydrobiological Studies, 53 (3), 227-236.
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• Toutain, M., Belouard, N., Renault, D., Haubrock, P.J., Kurtul, I., Aksu, S., Emiroğlu, Ö., Kouba, A., Tarkan, A.S., Balzani, P., 2024. Assessing the role of non-native species and artificial water bodies on the trophic and functional niche of Mediterranean freshwater fish communities. Science of the Total Environment, 938.
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• Soto, I., Balzani, P., Carneiro, L., Cuthbert, R.N., Macêdo, R., Serhan Tarkan, A., Ahmed, D.A., Bang, A., Bacela-Spychalska, K., Bailey, S.A., Baudry, T., Ballesteros-Mejia, L., Bortolus, A., Briski, E., Britton, J.R., Buřič, M., Camacho-Cervantes, M., Cano-Barbacil, C., Copilaș-Ciocianu, D., Coughlan, N.E., Courtois, P., Csabai, Z., Dalu, T., De Santis, V., Dickey, J.W.E., Dimarco, R.D., Falk-Andersson, J., Fernandez, R.D., Florencio, M., Franco, A.C.S., García-Berthou, E., Giannetto, D., Glavendekic, M.M., Grabowski, M., Heringer, G., Herrera, I., Huang, W., Kamelamela, K.L., Kirichenko, N.I., Kouba, A., Kourantidou, M., Kurtul, I., Laufer, G., Lipták, B., Liu, C., López-López, E., Lozano, V., Mammola, S., Marchini, A., Meshkova, V., Milardi, M., Musolin, D.L., Nuñez, M.A., Oficialdegui, F.J., Patoka, J., Pattison, Z., Pincheira-Donoso, D., Piria, M., Probert, A.F., Rasmussen, J.J., Renault, D., Ribeiro, F., Rilov, G., Robinson, T.B., Sanchez, A.E., Schwindt, E., South, J., Stoett, P., Verreycken, H., Vilizzi, L., Wang, Y.J., Watari, Y., Wehi, P.M., Weiperth, A., Wiberg-Larsen, P., Yapıcı, S., Yoğurtçuoğlu, B., Zenni, R.D., Galil, B.S., Dick, J.T.A., Russell, J.C., Ricciardi, A., Simberloff, D., Bradshaw, C.J.A., Haubrock, P.J., 2024. Taming the terminological tempest in invasion science. Biological Reviews, 99 (4), 1357-1390.
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• Kurtul, I., Tarkan, A.S., Sarı, H.M., Haubrock, P.J., Soto, I., Aksu, S., Britton, J.R., 2024. Exploring invasiveness and versatility of used microhabitats of the globally invasive Gambusia holbrooki. Science of the Total Environment, 925.
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• Haubrock, P.J., Soto, I., Ahmed, D.A., Ansari, A.R., Tarkan, A.S., Kurtul, I., Macêdo, R.L., Lázaro-Lobo, A., Toutain, M., Parker, B., Błońska, D., Guareschi, S., Cano-Barbacil, C., Dominguez Almela, V., Andreou, D., Moyano, J., Akalın, S., Kaya, C., Bayçelebi, E., Yoğurtçuoğlu, B., Briski, E., Aksu, S., Emiroğlu, Ö., Mammola, S., De Santis, V., Kourantidou, M., Pincheira-Donoso, D., Britton, J.R., Kouba, A., Dolan, E.J., Kirichenko, N.I., García-Berthou, E., Renault, D., Fernandez, R.D., Yapıcı, S., Giannetto, D., Nuñez, M.A., Hudgins, E.J., Pergl, J., Milardi, M., Musolin, D.L., Cuthbert, R.N., 2024. Biological invasions are a population-level rather than a species-level phenomenon. Global Change Biology, 30 (5).
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• Kurtul, I., Kaya, C., Kaykaç, H., Ilhan, A., Düzbastilar, F.O., Tosunoğlu, Z., Sari, H.M., Balzani, P., Tarkan, A.S., Haubrock, P.J., 2024. How fish populations in Lake Bafa (Western Anatolia) respond to ecological shifts. Aquatic Conservation Marine and Freshwater Ecosystems, 34 (5).
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• Bayçelebi, E., Kaya, C., Kurtul, I., Turan, D., Haubrock, P.J., Soto, I., Aksu, S., Tarkan, A.S., 2024. Decadal-scale regional variability in monitoring efforts significantly influences fish diversity trends in the Euphrates and Tigris catchments, Türkiye. Aquatic Conservation Marine and Freshwater Ecosystems, 34 (5).
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• Tarkan, A.S., Bayçelebi, E., Giannetto, D., Özden, E.D., Yazlık, A., Emiroğlu, Ö., Aksu, S., Uludağ, A., Aksoy, N., Baytaşoğlu, H., Kaya, C., Mutlu, T., Kırankaya, Ş.G., Ergüden, D., Per, E., Üremiş, İ., Candan, O., Kekillioğlu, A., Yoğurtçuoğlu, B., Ekmekçi, F.G., Başak, E., Özkan, H., Kurtul, I., Innal, D., Killi, N., Yapıcı, S., Ayaz, D., Çiçek, K., Mol, O., Çınar, E., Yeğen, V., Angulo, E., Cuthbert, R.N., Soto, I., Courchamp, F., Haubrock, P.J., 2024. Economic costs of non-native species in Türkiye: A first national synthesis. Journal of Environmental Management, 358.
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• Kaya, C., Kurtul, I., Bayçelebi, E., İlhan, A., Sarı, H.M., 2024. Actual Distributions and Validity of Petroleuciscus spp., with the Range Extension and Length-Weight Relationship Data in Case of Petroleuciscus ninae. Turkish Journal of Fisheries and Aquatic Sciences, 24 (2).
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• Tarkan, A.S., Kurtul, I., Błońska, D., Robert Britton, J., Haubrock, P.J., 2024. Resolving the issues of translocated species in freshwater invasions. Neobiota, 93, 177-186.
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• Tarkan, A.S., Kurtul, I., Britton, J.R., 2024. Comparing the ecological consequences of globally invasive fishes versus their F1 hybrids in recreational fisheries. Neobiota, 95, 267-278.
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• Ion, M.C., Bloomer, C.C., Bărăscu, T.I., Oficialdegui, F.J., Shoobs, N.F., Williams, B.W., Scheers, K., Clavero, M., Grandjean, F., Collas, M., Baudry, T., Loughman, Z., Wright, J.J., Ruokonen, T.J., Chucholl, C., Guareschi, S., Koese, B., Banyai, Z.M., Hodson, J., Hurt, M., Kaldre, K., Lipták, B., Fetzner, J.W., Cancellario, T., Weiperth, A., Birzaks, J., Trichkova, T., Todorov, M., Balalaikins, M., Griffin, B., Petko, O.N., Acevedo-Alonso, A., Elía, G.D., Śliwińska, K., Alekhnovich, A., Choong, H., South, J., Whiterod, N., Zorić, K., Haase, P., Soto, I., Brady, D.J., Haubrock, P.J., Torres, P.J., Şadrin, D., Vlach, P., Kaya, C., Jung, S.W., Kim, J.Y., Vermeersch, X.H.C., Bonk, M., Guiaşu, R., Harlioğlu, M.M., Devlin, J., Kurtul, I., Błońska, D., Boets, P., Masigol, H., Cabe, P.R., Jussila, J., Vrålstad, T., Beresford, D.V., Reid, S.M., Patoka, J., Strand, D.A., Tarkan, A.S., Steen, F., Abeel, T., Harwood, M., Auer, S., Kelly, S., Giantsis, I.A., Maciaszek, R., Alvanou, M.V., Aksu, Ö., Hayes, D.M., Kawai, T., Tricarico, E., Chakandinakira, A., Barnett, Z.C., Kudor, Ş.G., Beda, A.E., Vîlcea, L., Mizeranschi, A.E., Neagul, M., Licz, A., Cotoarbă, A.D., Petrusek, A., Kouba, A., Taylor, C.A., Pârvulescu, L., 2024. World of Crayfish™: a web platform towards real-time global mapping of freshwater crayfish and their pathogens. Peerj, 12 (10).
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• Kaya, C., Kurtul, I., Aksu, İ.L., Oral, M., Freyhof, J., 2024. Oxynoemacheilus chaboras, a new loach species from the Euphrates drainage in Türkiye (Teleostei, Nemacheilidae). Zoosystematics and Evolution, 100 (2), 457-468.
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• Kaya, C., İmre, H.B., Kurtul, I., 2024. Is Garra rezai (Teleostei, Cyprinidae) a species known only from two widely disjunct areas in the Tigris drainage? Zoosystematics and Evolution, 100 (2), 349-356.
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• Tarkan, A.S., Haubrock, P.J., Aksu, S., Mol, O., Balzani, P., Emiroğlu, Ö., Köse, E., Kurtul, I., Başkurt, S., Çınar, E., Oztopcu-Vatan, P., 2023. Predicting the potential implications of perch (Perca fluviatilis) introductions to a biodiversity-rich lake using stable isotope analysis. Scientific Reports, 13 (1).
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• Kurtul, I., Kaya, C., Şen, A., Özcan, C., Ercivan, D., Demiral, K., Güngör, M., İlhan, A., Sarı, H.M., 2023. An ichthyofaunal amendment with the length-weight relations and condition factors of some endemic and invasive freshwater fishes from Western Anatolia. Oceanological and Hydrobiological Studies, 52 (4), 389-398.
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• Kurtul, I., Tarkan, A.S., Britton, J.R., 2023. Promoting the use of non-lethal sample collection for analysing the trophic relationships of inshore flatfish populations using stable isotope analysis. Estuarine Coastal and Shelf Science, 290.
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• Tarkan, A.S., Mol, O., Aksu, S., Köse, E., Kurtul, I., Başkurt, S., Haubrock, P.J., Balzani, P., Çınar, E., Britton, J.R., Oztopcu-Vatan, P., Emiroğlu, Ö., 2023. Phenotypic responses to piscivory in invasive gibel carp populations. Aquatic Sciences, 85 (3).
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• Kurtul, I., Tarkan, A.S., Britton, J.R., 2023. Inter-tissue variability in the stable isotope values of European perch (Perca fluviatilis) and pumpkinseed (Lepomis gibbosus). Knowledge and Management of Aquatic Ecosystems.
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• Kurtul, I., Tarkan, A.S., Sarı, H.M., Britton, J.R., 2022. Climatic and geographic variation as a driver of phenotypic divergence in reproductive characters and body sizes of invasive Gambusia holbrooki. Aquatic Sciences, 84 (2).
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• Vilizzi, L., Copp, G.H., Hill, J.E., Adamovich, B., Aislabie, L., Akin, D., Al-Faisal, A.J., Almeida, D., Azmai, M.N.A., Bakiu, R., Bellati, A., Bernier, R., Bies, J.M., Bilge, G., Branco, P., Bui, T.D., Canning-Clode, J., Cardoso Ramos, H.A., Castellanos-Galindo, G.A., Castro, N., Chaichana, R., Chainho, P., Chan, J., Cunico, A.M., Curd, A., Dangchana, P., Dashinov, D., Davison, P.I., de Camargo, M.P., Dodd, J.A., Durland Donahou, A.L., Edsman, L., Ekmekçi, F.G., Elphinstone-Davis, J., Erős, T., Evangelista, C., Fenwick, G., Ferincz, Á., Ferreira, T., Feunteun, E., Filiz, H., Forneck, S.C., Gajduchenko, H.S., Gama Monteiro, J., Gestoso, I., Giannetto, D., Gilles, A.S., Gizzi, F., Glamuzina, B., Glamuzina, L., Goldsmit, J., Gollasch, S., Goulletquer, P., Grabowska, J., Harmer, R., Haubrock, P.J., He, D., Hean, J.W., Herczeg, G., Howland, K.L., İlhan, A., Interesova, E., Jakubčinová, K., Jelmert, A., Johnsen, S.I., Kakareko, T., Kanongdate, K., Killi, N., Kim, J.E., Kırankaya, Ş.G., Kňazovická, D., Kopecký, O., Kostov, V., Koutsikos, N., Kozic, S., Kuljanishvili, T., Kumar, B., Kumar, L., Kurita, Y., Kurtul, I., Lazzaro, L., Lee, L., Lehtiniemi, M., Leonardi, G., Leuven, R.S.E.W., Li, S., Lipinskaya, T., Liu, F., Lloyd, L., Lorenzoni, M., Luna, S.A., Lyons, T.J., Magellan, K., Malmstrøm, M., Marchini, A., Marr, S.M., Masson, G., Masson, L., McKenzie, C.H., Memedemin, D., 2021. A global-scale screening of non-native aquatic organisms to identify potentially invasive species under current and future climate conditions. Science of the Total Environment, 788.
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• Tarkan, A.S., Tricarico, E., Vilizzi, L., Bİlge, G., Ekmekçİ, F.G., Filiz, H., Giannetto, D., İlhan, A., Kİllİ, N., Kirankaya, G., Koutsikos, N., Kozic, S., Kurtul, I., Lazzaro, L., Marchini, A., Occhipinti-Ambrogi, A., Perdikaris, C., Piria, M., Pompei, L., Sari, H., Smeti, E., Stasolla, G., Top, N., Tsiamis, K., Vardakas, L., Yapici, S., Yoğurtçuoğlu, B., Copp, G.H., 2021. Risk of invasiveness of non-native aquatic species in the eastern Mediterranean region under current and projected climate conditions. European Zoological Journal, 88 (1), 1130-1143.
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Preprints

• Tarkan, A.S., Kurtul, I., Blonska, D., Britton, J.R., Haubrock, P.. Resolving the issues of translocated native species in freshwater invasions.
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